Notes on Liverwort Specimens from the Philippine Cordilleras in the Botanical Herbarium of the UPLB Museum of Natural History (CAHUP)
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Oct 4, 2019
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Abstract
An account of liverwort specimens collected from the Philippine Cordilleras, particularly from the Cordillera Administrative Region (CAR), deposited and accessioned in the Botanical Herbarium of the University of the Philippines Los Baños Museum of Natural History (CAHUP) is provided. A total of 539 accessions from the region were recorded, comprising 23.89% of the overall liverwort and hornwort collection of CAHUP. This represents a total of 76 species distributed in 41 genera and 26 families. The families Lepidoziaceae with 179 accessions, Plagiochilaceae with 100 accessions, and Herbertaceae with 48 accessions make up the majority of the collections from the region. The oldest dated specimen from the CAR deposited in the herbarium was collected by H.H. Bartlett in 1935 and the bulk of accessioned specimens from the region were collected in the 1970s by M. Alvarez with A. Tandang and Saprid from Mt. Data (Mountain Province) and Mt. Pulag (Benguet Province), respectively. The geographic extent of liverwort collection in the CAR, as reflected by the accessioned specimens, is restricted to few localities in the provinces of Benguet, Ifugao, and Mountain Province. Locality information and distribution records for noteworthy species are reported, with 20 species new to the CAR; new localities are reported for another 18 species. With the current state of liverwort collection and research in the CAR, the processing and accessioning of approximately 15,000 bryophyte specimens from different parts of the country, conducting inventories and examination of herbarium specimens in other herbaria, and carrying out extensive and systematic collection work in areas of high conservation value are expected to contribute more information on the ecology, richness and diversity, and distribution of liverworts in the region.
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Bakalin, V.A. (2016). A revision of Lepidoziaceae (Hepaticae) in the Russian Far East I. Bazzania. Botanica Pacifica 5(1): 33-52. doi: 10.17581/bp. 2016.05108
Boakes, E.H., McGowan, P.J.K., Fuller, R.A., Chang-qing, D., Clark, N.E., O'Connor, K. & Mace, G.M. (2010). Distorted views of biodiversity: Spatial and temporal bias in species occurrence data. PLoS Biology 8 e1000385. doi: 10.1371/journal.pbio.1000385
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Crandall-Stotler, B.J., Stotler, R.E., & Long D.G. (2009). Phylogeny and classification of the Marchantiophyta. Edinburgh Journal of Botany 66 (1): 155–198. doi: 10.1017/S096042860900 5393
Fernando, E.S., Suh, M.H., Lee, J.H., & Lee, D.K. (2008). Forest formations of the Philippines. Seoul: ASEAN-Korea Environmental Cooperation Unit.
Fernando, E.S. & Cereno, R.P. (2010). Biodiversity and natural resources management in the Mt. Pulag National Park, Philippines. In: Lapitan, P.G., E.S. Fernando, M.H. Suh, R.U. Fuentes, Y.K. Shin, N.M. Pampolina, M.L. Castillo, R.P. Cereno, J.H. Lee, S. Han, T.B. Choi, & D.K. Lee. Biodiversity and natural resources conservation in protected areas of Korea and the Philippines. Seoul: ASEAN-Korea Environmental Cooperation Unit.
Gapud, V.F. & Tan, B.C. (1982). The role of systematics in basic and applied biology. Unpublished manuscript.
Grolle, R. (1968a). Monographie der Gattung Nowellia. Journal of the Hattori Botanical Laboratory 31: 20-49.
Grolle, R. (1968b). Gottschelia – eine neue Jungermanniales-Gattung der Paläotropis. Journal of the Hattori Botanical Laboratory 31: 13-19.
Hattori, S. (1975). Studies on Asiatic species of Porella (Hepaticae). V. Journal of the Hattori Botanical Laboratory 39: 269-276.
Inoue, H. & Miller, H.A. (1965). Hepaticae from Kusiae, Caroline Islands. Bulletin of the National Science Museum [Tokyo] 8: 139-160.
Inoue, H. (1965). A small collection of liverworts from Isl. Batan, the Philippines. Hikobia 4: 272-276.
Inoue, H. (1974). Illustrations of Japanese Hepaticae. Tokyo. 193 pp.
Jack, J.B. & Stephani, F. (1892). Hepaticae Wallisianae. Hedwigia 31: 11-27.
Juslén, A. (2006). Revision of Asian Herbertus (Herbertaceae, Marchantiophyta). Annales Botanici Fennici 43: 409-436.
Katagiri, T., Sadamitsu, A., Miyauchi, H., Tsubota, H., & Deguchi, H. (2013). Taxonomic studies of the Trichocoleaceae of Southeast Asia. III. The genus Trichocolea Dumort. Hattoria 4: 1-42.
Khotimperwati, L., Kasiamdari, R.S., Santosa, & Daryono, B.S. (2018). Bazzania Gray
(Lepidoziaceae, Marchantiophyta) in Central Java, Indonesia. Biodiversitas 19(3): 875-887.
Kitagawa, N. (1973). Miscellaneous notes on little-known species of Hepaticae, 1-25. Journal of the Hattori Botanical Laboratory 36: 444-454.
Lee, G.E. (2013). A systematic revision of the genus Lejeunea Lib. (Marchantiophyta: Lejeuneaceae) in Malaysia. Cryptogamie, Bryologie 34(4): 381–484. doi: 10.7872/cryb.v34.iss4.2013.381
Lehmann, J.G.C. (1833). Novarum et minus cognitorum stirpium pugillus quintus. Hamburg: Meissner. 72 pp. doi: 10.5962/bhl.title.45011
Long, D.G. & J. Váňa. (2007). The genus Gottschelia Grolle (Jungermanniopsida, Lophoziaceae) in China, with a description of G. grollei, sp. nov. Journal of Bryology 29: 165-168.
Magurran, A.E., Baillie, S.R., Buckland, S.T., Dick, J.M., Elston, D.A., Scott, E.M., Smith, R.I., Somerfield, P.J. & Watt, A.D. (2010). Long term datasets in biodiversity research and monitoring: Assessing change in ecological communities through time. Trends in Ecology and Evolution 25: 574-582. doi: 10.1016/j.tree.2010.06.016
Malabrigo, P.L. Jr. (2013). Vascular flora of the tropical montane forest in Balbalasang-Balbalan National Park, Kalinga Province, Northern Luzon, Philippines. Asian Journal of Biodiversity 4: 1-22.
Miller, H.A., Whittier, H.O. & Whittier, B.A. (1983). Prodromus florae hepaticarum Polynesiae. Catalogue of Hepaticae and Anthocerotae. Bryophytorum Bibliotheca 25: 1-423.
Miller-Rushing, A.J., Primack, R.B., Primack, D., & Mukunda, S. (2006). Photographs and herbarium specimens as tools to document phenological changes in response to global warming. American Journal of Botany 93: 1667-1674.
O'Connell, A.F. Jr., Gilbert, A.T., & Hatfield, J.S. (2004). Contribution of natural history collection data to biodiversity assessment in national parks. Conservation Biology 18(5): 1254-1261.
Piippo, S. (1985). Bryophyte flora of Huon Peninsula, Papua New Guinea. X. Jackiellaceae, Scapaniaceae, Arnelliaceae and Acrobolbaceae.
Pócs, T. & Ninh, T. (2005). Contribution to the bryoflora of Vietnam, VI. On the liverwort flora of Vu Quang Nature Reserve. Acta Botanica Hungarica 47(1-2): 151-171.
Pócs, T. (2011). East African bryophytes XXIX. The Ceratolejeunea (Lejeuneaceae) species in the Indian Ocean islands. Polish Botanical Journal 56(2): 131-153.
Pócs, T., Brown, E.A., Cairns, A., Cargill, D.C. & Pócs, S. (2012). Contributions to the bryoflora of Australia, III. The genus Nowellia Mitt. (Cephaloziaceae, Jungermanniopsida). Acta Biologica Plantarum Agriensis 2: 21-26.
Ponder, W.F., Carter, G.A., Flemons, P. & Chapman, R.R. (2001). Evaluation of museum collection data for use in biodiversity assessment. Conservation Biology 15(3): 648-657.
Pyke, G.H. & Ehrlich, P.R. (2010). Biological collections and ecological/environmental research: A review, some observations and a look to the future. Biological Reviews 85: 247-266. doi: 10.1111/j.1469-185X.2009.00098.x
Robbirt, K.M., Davy, A.J., Hutchings, M.J., & Roberts, D.R. (2011). Validation of biological collections as a source of phenological data for use in climate change studies: a case study with the orchid Ophrys sphegodes. Journal of Ecology 99: 235-241. doi: 10.1111/j.13652745.2010.01727.x
Sastre-De Jesus, I. & Tan, B.C. (1995). Problems of bryophyte conservation in the tropics: A discussion, with case examples from Puerto Rico and the Philippines. Caribbean Journal of Science 31: 200–206.
Shaffer, H.B., Fisher, R.N. & Davidson, C. (1998). The role of natural history collections in documenting species declines. TREE 13(1): 2730.
So, M.L. & Grolle, R. (2000). Checklist of Plagiochila (Hepaticae) in Asia. Journal of the Hattori Botanical Laboratory 88: 199-243.
So, M.L. (2000). Plagiochila sect. Plagiochila (Hepaticae) in SE Asia and Melanesia,
with descriptions of two new species. New Zealand Journal of Botany 38(3): 425-432. doi: 10.1080/0028825X.2000.9512694
So, M.L. (2001a). On Plagiochila section Cobanae Carl in Asia and Melanesia. Cryptogamie, Bryologie 22(3): 179-186.
So, M.L. (2001b). Plagiochila (Hepaticae, Plagiochilaceae) in China. Systematic Botany Monographs 60: 1-214.
So, M.L. (2003a). The genus Schistochila in Asia. Journal of the Hattori Botanical Laboratory 93: 79-100.
So, M.L. (2003b). The genus Metzgeria (Hepaticae) in Asia. Journal of the Hattori Botanical Laboratory 94: 159-177.
Söderström, L., Hagborg, A., von Konrat, M., Bartholomew-Began, S., Bell, D., Briscoe, L., Brown, E., Cargill, D.C., Costa, D.P., CrandallStotler, B.J., Cooper, E.D., Dauphin, G., Engel, J.J., Feldberg, K., Glenny, D., Gradstein, S.R., He, X., Heinrichs, J., Hentschel, J., Ilkiu-Borges, A.L., Katagiri, T., Konstantinova, N.A., Larraín, J., Long, D.G., Nebel, M., Pócs, T., Felisa Puche, F., Reiner-Drehwald, E., Renner, M.A.M., SassGyarmati, A., Schäfer-Verwimp, A., Moragues, J.G.S., Stotler, R.E., Sukkharak, P., Thiers, B.M., Uribe, J., Váňa, J., Villarreal, J.C., Wigginton, M., Zhang, L., & Zhu, R.-L. (2016). World checklist of hornworts and liverworts. PhytoKeys 59: 1-828. doi: 10.3897/phytokeys.59.6261
Sukkharak, P. & Gradstein, S.R. (2014). A taxonomic revision of the genus Mastigolejeunea (Marchantiophyta: Lejeuneaceae). Nova Hedwigia 99(3/4): 279–345. doi: 10.1127/00295035/2014/0206
Sukkharak, P. (2015). A systematic monograph of the genus Thysananthus (Lejeuneaceae, Marchantiophyta). Phytotaxa 193(1): 1–81. doi: 10.11646/phytotaxa.193.1.1
Tan, B.C. & Engel, J.J. (1986). An annotated checklist of Philippine Hepaticae. Journal of the Hattori Botanical Laboratory 60: 283-355.
Tan, B.C. & Iwatsuki, Z. (1999). Four hot spots of moss diversity in Malesia. Bryobrothera 5: 247-252.
Tan, B.C. (2000). East and Southeast Asia. In: Hallingbäck, T. & N. Hodgetts (compilers). Mosses, liverworts, and hornworts. Status survey and conservation action plan for bryophytes. IUCN/SSC Bryophyte Specialist Group. Gland, Switzerland and Cambridge, United Kingdom: International Union for the Conservation of Nature and Natural Resources.
Thiers, B. (2015 [continuously updated]). Index Herbariorum: A global directory of public herbaria and associated staff. New York Botanical Garden’s Virtual Herbarium. Available from http://sweetgum.nybg.org/science/ih/. Last accessed 22 September 2019.
Váňa, J. & Long, D.G. (2009). Jungermanniaceae of the Sino-Himalayan region. Nova Hedwigia 89(34): 485-517.
Váňa, J. & Piippo, S. (1989). Bryophyte flora of Huon Peninsula, Papua New Guinea. XXXI. Cephaloziaceae subfam. Alobielloideae, Cephaloziellaceae, Antheliaceae and Lophoziaceae. Annales Botanici Fennici 26(3): 263-290.
Váňa, J. (1992). The bryophytes of Sabah (North Borneo) with special reference to the BRYOTROP transect of Mount Kinabalu. XVI. Cephaloziellaceae and Jackiellaceae (Hepaticae). Willdenowia 22(1/2): 167-169.
Ward, D.F. (2012). More than just records: Analysing natural history collections for biodiversity planning. PLoS ONE 7(11): e50346. doi: 10.1371/journal.pone.0050346
Zhu, R.L., Shu, L., He, Q., & Wei, Y.M. (2018). Soella (Marchantiophyta: Lejeuneaceae), a new genus from China and Japan. The Bryologist 121(3): 324-339. doi: 10.1639/0007-2745-121.3.324
Zhu, R.-L., So, M.L. & Wang, Y.F. (2002). The genus Cheilolejeunea (Hepaticae, Lejeuneaceae) in China. Nova Hedwigia 75(3-4): 387-408. doi: 10.1127/0029-5035/2002/0075-0387
Aryanti, N.S. & Gradstein, S.R. (2007). Wallace’s line and the distribution of the liverworts of Sulawesi. Cryptogamie, Bryologie 28(1): 3-14
Bakalin, V.A. (2013). New taxa of Solenostoma and Plectocolea and other taxonomic novelties based on study of collections in the New York Botanical Garden Herbarium. Polish Botanical Journal 58(1): 127-142. doi: 10.2478/pbj-2013-0014
Bakalin, V.A. (2016). A revision of Lepidoziaceae (Hepaticae) in the Russian Far East I. Bazzania. Botanica Pacifica 5(1): 33-52. doi: 10.17581/bp. 2016.05108
Boakes, E.H., McGowan, P.J.K., Fuller, R.A., Chang-qing, D., Clark, N.E., O'Connor, K. & Mace, G.M. (2010). Distorted views of biodiversity: Spatial and temporal bias in species occurrence data. PLoS Biology 8 e1000385. doi: 10.1371/journal.pbio.1000385
Bonner, C.E.B. (1963). Index Hepaticarum. Pars IV. Ceratolejeunea to Cystolejeunea. Weinheim: J. Cramer. pp. 637-926.
Conservation International - Department of Environment and Natural Resources-Protected Areas and Wildlife Bureau – Haribon Foundation [CI-DENR-PAWB-Haribon]. (2006). Priority sites for conservation in the Philippines: Key Biodiversity Areas. Quezon City, Philippines. 24 pp.
Crandall-Stotler, B.J., Stotler, R.E., & Long D.G. (2009). Phylogeny and classification of the Marchantiophyta. Edinburgh Journal of Botany 66 (1): 155–198. doi: 10.1017/S096042860900 5393
Fernando, E.S., Suh, M.H., Lee, J.H., & Lee, D.K. (2008). Forest formations of the Philippines. Seoul: ASEAN-Korea Environmental Cooperation Unit.
Fernando, E.S. & Cereno, R.P. (2010). Biodiversity and natural resources management in the Mt. Pulag National Park, Philippines. In: Lapitan, P.G., E.S. Fernando, M.H. Suh, R.U. Fuentes, Y.K. Shin, N.M. Pampolina, M.L. Castillo, R.P. Cereno, J.H. Lee, S. Han, T.B. Choi, & D.K. Lee. Biodiversity and natural resources conservation in protected areas of Korea and the Philippines. Seoul: ASEAN-Korea Environmental Cooperation Unit.
Gapud, V.F. & Tan, B.C. (1982). The role of systematics in basic and applied biology. Unpublished manuscript.
Grolle, R. (1968a). Monographie der Gattung Nowellia. Journal of the Hattori Botanical Laboratory 31: 20-49.
Grolle, R. (1968b). Gottschelia – eine neue Jungermanniales-Gattung der Paläotropis. Journal of the Hattori Botanical Laboratory 31: 13-19.
Hattori, S. (1975). Studies on Asiatic species of Porella (Hepaticae). V. Journal of the Hattori Botanical Laboratory 39: 269-276.
Inoue, H. & Miller, H.A. (1965). Hepaticae from Kusiae, Caroline Islands. Bulletin of the National Science Museum [Tokyo] 8: 139-160.
Inoue, H. (1965). A small collection of liverworts from Isl. Batan, the Philippines. Hikobia 4: 272-276.
Inoue, H. (1974). Illustrations of Japanese Hepaticae. Tokyo. 193 pp.
Jack, J.B. & Stephani, F. (1892). Hepaticae Wallisianae. Hedwigia 31: 11-27.
Juslén, A. (2006). Revision of Asian Herbertus (Herbertaceae, Marchantiophyta). Annales Botanici Fennici 43: 409-436.
Katagiri, T., Sadamitsu, A., Miyauchi, H., Tsubota, H., & Deguchi, H. (2013). Taxonomic studies of the Trichocoleaceae of Southeast Asia. III. The genus Trichocolea Dumort. Hattoria 4: 1-42.
Khotimperwati, L., Kasiamdari, R.S., Santosa, & Daryono, B.S. (2018). Bazzania Gray
(Lepidoziaceae, Marchantiophyta) in Central Java, Indonesia. Biodiversitas 19(3): 875-887.
Kitagawa, N. (1973). Miscellaneous notes on little-known species of Hepaticae, 1-25. Journal of the Hattori Botanical Laboratory 36: 444-454.
Lee, G.E. (2013). A systematic revision of the genus Lejeunea Lib. (Marchantiophyta: Lejeuneaceae) in Malaysia. Cryptogamie, Bryologie 34(4): 381–484. doi: 10.7872/cryb.v34.iss4.2013.381
Lehmann, J.G.C. (1833). Novarum et minus cognitorum stirpium pugillus quintus. Hamburg: Meissner. 72 pp. doi: 10.5962/bhl.title.45011
Long, D.G. & J. Váňa. (2007). The genus Gottschelia Grolle (Jungermanniopsida, Lophoziaceae) in China, with a description of G. grollei, sp. nov. Journal of Bryology 29: 165-168.
Magurran, A.E., Baillie, S.R., Buckland, S.T., Dick, J.M., Elston, D.A., Scott, E.M., Smith, R.I., Somerfield, P.J. & Watt, A.D. (2010). Long term datasets in biodiversity research and monitoring: Assessing change in ecological communities through time. Trends in Ecology and Evolution 25: 574-582. doi: 10.1016/j.tree.2010.06.016
Malabrigo, P.L. Jr. (2013). Vascular flora of the tropical montane forest in Balbalasang-Balbalan National Park, Kalinga Province, Northern Luzon, Philippines. Asian Journal of Biodiversity 4: 1-22.
Miller, H.A., Whittier, H.O. & Whittier, B.A. (1983). Prodromus florae hepaticarum Polynesiae. Catalogue of Hepaticae and Anthocerotae. Bryophytorum Bibliotheca 25: 1-423.
Miller-Rushing, A.J., Primack, R.B., Primack, D., & Mukunda, S. (2006). Photographs and herbarium specimens as tools to document phenological changes in response to global warming. American Journal of Botany 93: 1667-1674.
O'Connell, A.F. Jr., Gilbert, A.T., & Hatfield, J.S. (2004). Contribution of natural history collection data to biodiversity assessment in national parks. Conservation Biology 18(5): 1254-1261.
Piippo, S. (1985). Bryophyte flora of Huon Peninsula, Papua New Guinea. X. Jackiellaceae, Scapaniaceae, Arnelliaceae and Acrobolbaceae.
Pócs, T. & Ninh, T. (2005). Contribution to the bryoflora of Vietnam, VI. On the liverwort flora of Vu Quang Nature Reserve. Acta Botanica Hungarica 47(1-2): 151-171.
Pócs, T. (2011). East African bryophytes XXIX. The Ceratolejeunea (Lejeuneaceae) species in the Indian Ocean islands. Polish Botanical Journal 56(2): 131-153.
Pócs, T., Brown, E.A., Cairns, A., Cargill, D.C. & Pócs, S. (2012). Contributions to the bryoflora of Australia, III. The genus Nowellia Mitt. (Cephaloziaceae, Jungermanniopsida). Acta Biologica Plantarum Agriensis 2: 21-26.
Ponder, W.F., Carter, G.A., Flemons, P. & Chapman, R.R. (2001). Evaluation of museum collection data for use in biodiversity assessment. Conservation Biology 15(3): 648-657.
Pyke, G.H. & Ehrlich, P.R. (2010). Biological collections and ecological/environmental research: A review, some observations and a look to the future. Biological Reviews 85: 247-266. doi: 10.1111/j.1469-185X.2009.00098.x
Robbirt, K.M., Davy, A.J., Hutchings, M.J., & Roberts, D.R. (2011). Validation of biological collections as a source of phenological data for use in climate change studies: a case study with the orchid Ophrys sphegodes. Journal of Ecology 99: 235-241. doi: 10.1111/j.13652745.2010.01727.x
Sastre-De Jesus, I. & Tan, B.C. (1995). Problems of bryophyte conservation in the tropics: A discussion, with case examples from Puerto Rico and the Philippines. Caribbean Journal of Science 31: 200–206.
Shaffer, H.B., Fisher, R.N. & Davidson, C. (1998). The role of natural history collections in documenting species declines. TREE 13(1): 2730.
So, M.L. & Grolle, R. (2000). Checklist of Plagiochila (Hepaticae) in Asia. Journal of the Hattori Botanical Laboratory 88: 199-243.
So, M.L. (2000). Plagiochila sect. Plagiochila (Hepaticae) in SE Asia and Melanesia,
with descriptions of two new species. New Zealand Journal of Botany 38(3): 425-432. doi: 10.1080/0028825X.2000.9512694
So, M.L. (2001a). On Plagiochila section Cobanae Carl in Asia and Melanesia. Cryptogamie, Bryologie 22(3): 179-186.
So, M.L. (2001b). Plagiochila (Hepaticae, Plagiochilaceae) in China. Systematic Botany Monographs 60: 1-214.
So, M.L. (2003a). The genus Schistochila in Asia. Journal of the Hattori Botanical Laboratory 93: 79-100.
So, M.L. (2003b). The genus Metzgeria (Hepaticae) in Asia. Journal of the Hattori Botanical Laboratory 94: 159-177.
Söderström, L., Hagborg, A., von Konrat, M., Bartholomew-Began, S., Bell, D., Briscoe, L., Brown, E., Cargill, D.C., Costa, D.P., CrandallStotler, B.J., Cooper, E.D., Dauphin, G., Engel, J.J., Feldberg, K., Glenny, D., Gradstein, S.R., He, X., Heinrichs, J., Hentschel, J., Ilkiu-Borges, A.L., Katagiri, T., Konstantinova, N.A., Larraín, J., Long, D.G., Nebel, M., Pócs, T., Felisa Puche, F., Reiner-Drehwald, E., Renner, M.A.M., SassGyarmati, A., Schäfer-Verwimp, A., Moragues, J.G.S., Stotler, R.E., Sukkharak, P., Thiers, B.M., Uribe, J., Váňa, J., Villarreal, J.C., Wigginton, M., Zhang, L., & Zhu, R.-L. (2016). World checklist of hornworts and liverworts. PhytoKeys 59: 1-828. doi: 10.3897/phytokeys.59.6261
Sukkharak, P. & Gradstein, S.R. (2014). A taxonomic revision of the genus Mastigolejeunea (Marchantiophyta: Lejeuneaceae). Nova Hedwigia 99(3/4): 279–345. doi: 10.1127/00295035/2014/0206
Sukkharak, P. (2015). A systematic monograph of the genus Thysananthus (Lejeuneaceae, Marchantiophyta). Phytotaxa 193(1): 1–81. doi: 10.11646/phytotaxa.193.1.1
Tan, B.C. & Engel, J.J. (1986). An annotated checklist of Philippine Hepaticae. Journal of the Hattori Botanical Laboratory 60: 283-355.
Tan, B.C. & Iwatsuki, Z. (1999). Four hot spots of moss diversity in Malesia. Bryobrothera 5: 247-252.
Tan, B.C. (2000). East and Southeast Asia. In: Hallingbäck, T. & N. Hodgetts (compilers). Mosses, liverworts, and hornworts. Status survey and conservation action plan for bryophytes. IUCN/SSC Bryophyte Specialist Group. Gland, Switzerland and Cambridge, United Kingdom: International Union for the Conservation of Nature and Natural Resources.
Thiers, B. (2015 [continuously updated]). Index Herbariorum: A global directory of public herbaria and associated staff. New York Botanical Garden’s Virtual Herbarium. Available from http://sweetgum.nybg.org/science/ih/. Last accessed 22 September 2019.
Váňa, J. & Long, D.G. (2009). Jungermanniaceae of the Sino-Himalayan region. Nova Hedwigia 89(34): 485-517.
Váňa, J. & Piippo, S. (1989). Bryophyte flora of Huon Peninsula, Papua New Guinea. XXXI. Cephaloziaceae subfam. Alobielloideae, Cephaloziellaceae, Antheliaceae and Lophoziaceae. Annales Botanici Fennici 26(3): 263-290.
Váňa, J. (1992). The bryophytes of Sabah (North Borneo) with special reference to the BRYOTROP transect of Mount Kinabalu. XVI. Cephaloziellaceae and Jackiellaceae (Hepaticae). Willdenowia 22(1/2): 167-169.
Ward, D.F. (2012). More than just records: Analysing natural history collections for biodiversity planning. PLoS ONE 7(11): e50346. doi: 10.1371/journal.pone.0050346
Zhu, R.L., Shu, L., He, Q., & Wei, Y.M. (2018). Soella (Marchantiophyta: Lejeuneaceae), a new genus from China and Japan. The Bryologist 121(3): 324-339. doi: 10.1639/0007-2745-121.3.324
Zhu, R.-L., So, M.L. & Wang, Y.F. (2002). The genus Cheilolejeunea (Hepaticae, Lejeuneaceae) in China. Nova Hedwigia 75(3-4): 387-408. doi: 10.1127/0029-5035/2002/0075-0387